Humphries, Julian & R.R. Miller. 1981. "A remarkable species flock of pupfishes, genus Cyprinodon, from Yucatán, Mexico". Copeia. 1981(1):52-64 (ffm00180) (resumen)
Humphries, Julian. 1984. "Cyprinodon verecundus, n. sp., a fifth species of pupfish from Laguna Chichancanab". Copeia. 1984(1):58-68 (ffm00193)