Doadrio, Ignacio & J.A. Carmona, E. Martínez, A. de Sostoa. 1999. "Genetic variation and taxonomic analysis of the subgenus Profundulus". Journal of Fish Biology. 55(4):751-766 (ffm00215) (resumen)
Doadrio, Ignacio & O. Domínguez-Domínguez. 2004. "Phylogenetic relationships within the fish family Goodeidae based on cytochrome b sequence data". Molecular Phylogenetics and Evolution. (31):416-430 (ffm00620) (resumen)