最初的描述(命名) :
- Girard, Charles Frederic. 1854. "Descriptions of new fishes, collected by Dr. A. L. Heermann, naturalist attached to the survey of the Pacific railroad route, under Lieut. R. S. Williamson, U. S. A". Proceedings of the Academy of Natural Sciences of Philadelphia. 7:129-140 (ffm00543)