最初的描述(命名) :
- Jordan, David Starr & B.W. Evermann. 1896. "The fishes of North and Middle America: A descriptive catalogue of the species of fishes and fish-like vertebrates found in the waters of North America, north of the Isthmus of Panama. Part I". Behavioural Processes. 47:1-1240 - (ffm00055)